• XU Kewang

  XU Kewang

  1 State Key Laboratory of Biocontrol, Guangdong Provincial Key Laboratory of Plant Resources, and School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China
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• LIAO Wenbo

  LIAO Wenbo

  1 State Key Laboratory of Biocontrol, Guangdong Provincial Key Laboratory of Plant Resources, and School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China
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• JIN Jianhua

  JIN Jianhua

  1 State Key Laboratory of Biocontrol, Guangdong Provincial Key Laboratory of Plant Resources, and School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China 2 State Key Laboratory of Palaeobiology and Stratigraphy (Nanjing Institute of Geology and Palaeontology, CAS), Nanjing 210008, China
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• LIU Xiaoyan

  LIU Xiaoyan

  1 State Key Laboratory of Biocontrol, Guangdong Provincial Key Laboratory of Plant Resources, and School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China 2 State Key Laboratory of Palaeobiology and Stratigraphy (Nanjing Institute of Geology and Palaeontology, CAS), Nanjing 210008, China
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This study was supported by the National Natural Science Foundation of China (Grant No. 41210001), the joint Project of the National Natural Science Foundation of China and the Russian Foundation for Basic Research (Grant Nos. 41611130044, 16-55-53007), the Fundamental Research Funds for the Central Universities (Grant no. 16lgjc28), State Key Laboratory of Palaeobiology and Stratigraphy (Nanjing Institute of Geology and Palaeontology, CAS) (No.163109), and the Scientific Research Fund, Hongda Zhang, Sun Yat-sen University. We sincerely thank Xianqiu Zhang for collecting the fossils. We also offer sincere gratitude to Qingqing Xu, Meng Han for their constructive suggestions on this study, and Pavel Alexeev, researcher of Komarov Botanical Institute for providing references. We are grateful to Prof. Robert A. Spicer (The Open University, UK) for English improvements.