1 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China；2 Henan Geological Museum, Zhengzhou 450016, Henan Province, China
The first author thanks Drs. D. Winkler (SMU, USA) and D. Hone (University College, Ireland) for their helpful and constructive comments on the earlier versions of this paper, Y. Q. Zhang (CAGS) prepared the specimens. This research was supported by grants from the Natural Science Foundation of China (90914003; 40872017), and the Basic Outlay of Scientific Research Work from the Ministry of Science and Technology (2011-SYS-02).
A new pterosaur Moganopterus zhuiana gen. et sp. nov. is erected based on a complete skull with lower jaws and anterior cervical vertebrae. It is characterized by much elongated upper and lower jaws with at least 62 total, long, curved teeth with sharp tips, a well developed parietal crest extending posterodorsally, forming an angle of 15 degrees with the ventral margin of the skull, the ratio of length to width of cervical vertebrae greater than 5:1. The skull length is 750 mm, and it is the largest toothed pterosaur found so far in the world. Based on this new pterosaur, the Boreopteridae can be divided into two subgroups: Boreopterinae sub-fam. nov. and Moganopterinae sub-fam. nov., which is also confirmed by the phylogenetic analysis.