• WU Wenhao

  WU Wenhao

  1 Research Center of Palaeontology & Stratigraphy, College of Earth Science, Jilin University, Changchun 130026, China
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• DONG Nan

  DONG Nan

  2 College of Earth Science, Jilin University, Changchun 130061, China
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• LI Xiaobo

  LI Xiaobo

  2 College of Earth Science, Jilin University, Changchun 130061, China 3 Dinosaur Evolution Research Centre, College of Earth Science, Jilin University, Changchun 130061, China
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• ZHOU Changfu

  ZHOU Changfu

  4 College of Earth Science and Engineering, Shandong University of Science and Technology, Qingdao, Shandong 266590, China
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• FENG Nan

  FENG Nan

  5 Heilongjiang Institute of Geological Sciences, Harbin 150080, China
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• Robert R. REISZ

  Robert R. REISZ

  3 Dinosaur Evolution Research Centre, College of Earth Science, Jilin University, Changchun 130061, China 6 Department of Biology, University of Toronto Mississauga, Mississauga, ON L5L 1C6, Canada
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This study was supported by the National Science Foundation of China (Grant No. 31670215), Special Program of Key Lab for Evolution of Past Life and Environment in Northeast Asia (Jilin University), Ministry of Education, China (Grant No. 201801).