• Alberto VALENCIANO

  Alberto VALENCIANO

  1 Research and Exhibitions Department, Iziko Museums of South Africa, PO Box 61, Cape Town 8000, South Africa 2 Department of Biological Sciences, University of Cape Town, Palaeobiological Research Group, Private Bag X3, Rhodes Gift 7701, Cape Town, South Africa
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• JIANGZUO Qigao

  JIANGZUO Qigao

  3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing100044, China 4 CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 5 University of Chinese Academy of Sciences, Beijing 100049, China 6 Department of Paleontology, American Museum of Natural History, USA
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• WANG Shiqi

  WANG Shiqi

  3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing100044, China 4 CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China；7 CAS Center for Excellence in Tibetan Plateau Earth Sciences Beijing 100101, China
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• LI Chunxiao

  LI Chunxiao

  3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing100044, China 4 CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 5 University of Chinese Academy of Sciences, Beijing 100049, China
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• ZHANG Xiaoxiao

  ZHANG Xiaoxiao

  3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing100044, China 4 CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 5 University of Chinese Academy of Sciences, Beijing 100049, China
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• YE Je

  YE Je

  3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing100044, China 4 CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 5 University of Chinese Academy of Sciences, Beijing 100049, China 6 Department of Paleontology, American Museum of Natural History, USA；8 Tianjin Natural History Museum, Tianjin, 300201, China
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The current work was supported by the Chinese Academy of Sciences (Grant Nos.XDA20070203, XDB26000000, QYZDY-SSW-DQC-22, GJHZ1885), the National Natural Science Foundation of China (Grant Nos. 41430102 and 41772018, 41625005). The support of the DST-NFR Centre of Excellence in Palaeosciences (CoE-Pal) toward this research for A. Valenciano (COE2018-09POST) is hereby acknowledged.