• XING Lida

  XING Lida

  1 State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China；2 School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China
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• Ryan C. MCKELLAR

  Ryan C. MCKELLAR

  3 Royal Saskatchewan Museum, Regina, Saskatchewan S4P 4W7, Canada；4 Biology Department, University of Regina, Regina, Saskatchewan S4S 0A2, Canada；5 Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66405, USA
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• GAO Zhizhong

  GAO Zhizhong

  6 Department of Biology, Xinzhou Teachers University, Xinzhou 034000, Shanxi, China
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This research was funded by the Natural Sciences and Engineering Research Council of Canada (2015-00681), the National Geographic Society, USA (EC0768-15); National Natural Science Foundation of China (No. 41772008); National Natural Science Foundation of China (No. 31872198).