• LI Yunfeng

  LI Yunfeng

  1 Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Jilin University, Changchun 130026, China 2 Research Center of Palaeontology & Stratigraphy, Jilin University, Changchun 130026, Jilin, China
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• HUANG Wei

  HUANG Wei

  3 Jilin Normal University, Jilin, 136000, Jilin, China
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• WANG Hongshan

  WANG Hongshan

  4 Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA
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• David L. DILCHER

  David L. DILCHER

  1 Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Jilin University, Changchun 130026, China 2 Research Center of Palaeontology & Stratigraphy, Jilin University, Changchun 130026, Jilin, China；5 Department of Earth and Atmospheric Sciences, Indiana University, Bloomington, Indiana 47405, USA
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• TAN Xiao

  TAN Xiao

  1 Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Jilin University, Changchun 130026, China 2 Research Center of Palaeontology & Stratigraphy, Jilin University, Changchun 130026, Jilin, China；5 Department of Earth and Atmospheric Sciences, Indiana University, Bloomington, Indiana 47405, USA
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• LI Tao

  LI Tao

  1 Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Jilin University, Changchun 130026, China 2 Research Center of Palaeontology & Stratigraphy, Jilin University, Changchun 130026, Jilin, China
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• NA Yuling

  NA Yuling

  1 Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Jilin University, Changchun 130026, China 2 Research Center of Palaeontology & Stratigraphy, Jilin University, Changchun 130026, Jilin, China
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• SUN Chunlin

  SUN Chunlin

  1 Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Jilin University, Changchun 130026, China 2 Research Center of Palaeontology & Stratigraphy, Jilin University, Changchun 130026, Jilin, China
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This study was supported by the China Postdoctoral Science Foundation (grant No. 2018M631866), the National Natural Science Foundation of China (grant No. 41172009), China Geological Survey Grant (grant No. 1212011120149), Doctoral Fund of Ministry of Education of China (grant No. 20100061110034) and Project ‘‘111’’, China (grant No. B-06008).