The Late Carboniferous metazoan framework reefs are unusually well developed in Ziyun region, southern Guizhou Province, China, even in a global perspective. The main reefbuilding organisms include Formitchevella, Ivanovia cf. manchurica, phylloid algae, and “unnamed reefbuilding organisms”, and along with other inhabitants they constitute different reef communities. Within a given community, the organisms compete for space while maintaining symbiosis. In addition, ecological replacement also occurs among the different communities. Intraspecies competition drives Formitchevella skeleton to grow continuously, and sometimes the skeleton may be curled to avoid the spatial competition with the neighbors. As for interspecies competition, the “unnamed reefbuilding organisms” and Ivanovia cf. manchurica maintain an active and direct spatial competition through enwinding and contacting each other. Formitchevella and Ivanovia cf. manchurica keep a balanced competition and reach a competitive equilibrium, whereas phylloid algae surpass corals with their sheer number of countless individuals. The phylloid algae and the brachiopods have special ecological interactions, i.e., maintaining symbiosis while still competing for living space. In the reefbuilding process of Bianping village coral reef, the replacement of Formitchevella community and its lower communities resulted in the formation of Late Carboniferous largescale coral reef.